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Mathblock
\frac{da(t)}{dt} + \left[ \frac{1}{\tau_{act}} \cdot \Big( \beta + \left(1-\beta\right) u(t) \Big) \right] \cdot a(t) = \frac{u(t)}{\tau_{act}}

where 

Mathinline
body--uriencoded--\beta = \frac%7B\tau_%7Bact%7D%7D%7B\tau_%7Bdeact%7D%7D


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excerpt-include
S4.1 Excitation–activation dynamics
S4.1 Excitation–activation dynamics


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